Articles in international scientific journals with referee practice

(updated December 2012)

Laaksonen, T., Korpimäki, E. & Hakkarainen, H., 2002. Interactive effects of parental age and environmental variation on the breeding performance of Tengmalm’s owls. Journal of Animal Ecology 71, 23-31. PDF

Fargallo, J.A., Laaksonen, T., Pöyri, V., Korpimäki, E., 2002. Inter-sexual differences in the immune response of Eurasian kestrel nestlings under food shortage. Ecology Letters 5, 95-101. PDF

Fargallo, J.A., Laaksonen, T., Korpimäki, E., Pöyri, V., Griffith, S.C., Valkama, J., 2003. Size-mediated dominance and begging behaviour in Eurasian kestrel broods. Evolutionary Ecology Research 5, 549-558. PDF

Laaksonen, T., 2004. Hatching asynchrony as a bet-hedging strategy – an offspring diversity hypothesis. Oikos 104, 616-620. PDF

Laaksonen, T., Fargallo, JA., Korpimäki, E., Lyytinen, S., Valkama, J., Pöyri, V., 2004. Year- and sex-dependent effects of experimental brood sex ratio manipulation on fledging condition of Eurasian kestrels. Journal of Animal Ecology 73, 342-352. PDF

Laaksonen, T., Lyytinen, S., Korpimäki, E., 2004. Sex-specific recruitment and brood sex ratios of Eurasian kestrels in a seasonally and annually fluctuating environment. Evolutionary Ecology 18, 215-230. PDF

Ahola, M., Laaksonen, T., Sippola, K., Eeva, T., Rainio, K., Lehikoinen, E., 2004. Variation in climate warming along the migration route decouples arrival and breeding dates. Global Change Biology 10, 1610-1617. PDF.

Laaksonen, T., Hakkarainen, H., Korpimäki, E., 2004. Lifetime reproduction of a forest-dwelling owl increases with age and area of forests. Proceedings of the Royal Society of London B (suppl.) 271, S461-S464. PDF.

Rainio, K., Laaksonen, T., Ahola, M., Vähätalo, A.V., Lehikoinen, E., 2006. Climatic responses in spring migration of boreal and arctic birds in relation to wintering area and taxonomy. Journal of Avian Biology 37, 507-515. PDF.


Laaksonen, J., Laaksonen, T., Itämies, J., Rytkönen, S., Välimäki, P., 2006. A new efficient bait trap model for lepidoptera surveys – the “Oulu-Model”. Entomologica Fennica 17, 153-160. PDF


Laaksonen, T., Ahola, M., Eeva, T., Väisänen, R.A., Lehikoinen, E., 2006. Climate change, migratory connectivity and changes in laying date and clutch size of the pied flycatcher. Oikos 114, 277-290. PDF

Eeva, T., Hakkarainen, H., Laaksonen, T., Lehikoinen, E., 2006. Environmental pollution has sex-dependent effects on local survival. Biology Letters 2, 298-300. PDF

Fargallo, J.A., Laaksonen, T., Korpimäki, E., Wakamatsu, K., 2007. A melanin-based trait reflects environmental growth conditions of nestling male Eurasian kestrels. Evolutionary Ecology 21, 157-171. PDF

Dell’Arte, G.A., Laaksonen, T., Norrdahl, K., Korpimäki, E., 2007.  Variation in the diet composition of a generalist predator, the red fox, in relation to season and density of main prey. Acta Oecologica 31, 276-281. PDF

Ahola, M., Laaksonen, T., Eeva, T., Lehikoinen, E., 2007. Climate change can alter competitive relationships between resident and migratory birds. Journal of Animal Ecology 76, 1045-1052. PDF

Hakkarainen, H., Korpimäki, E., Laaksonen, T., Nikula, A., Suorsa, P. 2007. Survival of male Tengmalm’s owls increases with cover of old forest in the territory. Oecologia 155: 479-486. PDF

Helle, S., Laaksonen, T., Adamsson, A., Paranko, J., Huitu, O. 2008. Female field voles with high testosterone and glucose levels produce male-biased litters. Animal Behaviour 75: 1031-1039. PDF

Mäntylä, E., Klemola, T., Sirkiä, P., Laaksonen, T. 2008. Low light reflectance at the visible range may explain attraction of birds to defoliated trees. Behavioral Ecology, 19: 325-330. PDF

Laaksonen, T., Negro, J.J., Lyytinen, S., Valkama, J., Ots, I., Korpimäki, E. 2008.
Effects of experimental brood size manipulation and gender on carotenoid levels of Eurasian kestrels Falco tinnunculus. PLoS ONE 3: e2374. PDF

Korpimäki, E., Hakkarainen, H., Laaksonen, T. & Vasko, V. 2008. Responses of owls and Eurasian Kestrels to natural and human induced spatiotemporal variation. Scottish Birds 28: 19-27.

Eeva, T., Ahola, M., Laaksonen, T., Lehikoinen, E. 2008. The effects of sex, age and breeding success on breeding dispersal of pied flycatchers along a pollution gradient. Oecologia 157: 231-238. PDF

Mäntylä, E., Alessio, G.A., Blande, J.D., Heijari, J., Holopainen, J.K., Laaksonen, T., Piirtola, P., Klemola, T. 2008. From plants to birds: higher avian predation rates in trees responding to insect herbivory. PLoS ONE 3: e2832. PDF

Helle, S., Laaksonen, T. 2009. Latitudinal gradient in 2D:4D. Archives of Sexual Behaviour 38: 1-3. PDF

Alcaide, M., Serrano, D., Negro, J. J., Tella, J.L.Laaksonen, T., Müller, C., Gal, A., Korpimäki, E. 2009.
Population fragmentation leads to isolation by distance but not genetic impoverishment in the philopatric Lesser Kestrel: a comparison with the widespread and sympatric Eurasian Kestrel. Heredity 102: 190-198.

Pitala, N., Ruuskanen, S., Laaksonen, T., Doligez, B., Tschirren, B., Gustafsson,
L. 2009. The effects of experimentally manipulated yolk androgens on growth and immune function of male and female nestling collared flycatchers. Journal of Avian Biology 40: 225-230.

Lehtonen, P., Primmer, CR, Laaksonen, T.
2009. Different traits affect gain of extra-pair paternity and loss of paternity in the pied flycatcher (Ficedula hypoleuca). Animal Behaviour 77: 1103-1110.

Ruuskanen, S., Doligez, B., Tschirren, B., Pitala, N., Gustafsson, L., Groothuis, T.G.G., Laaksonen, T. 2009. Yolk androgens do not appear to mediate sexual conflict over parental investment in the collared flycatcher Ficedula albicollis. Hormones and Behaviour 55: 514-519.

Lilley, T., Laaksonen, T., Huitu, O. & Helle, S. 2009. Digit length ratio (2D/4D): comparing measurements from x-rays and photographs in field voles (Microtus agrestis). Behavioral Ecology and Sociobiology 63: 1539-1547.


Korpimäki, E., Hakkarainen, H., Laaksonen, T. & Vasko, V. 2009. Responses of owls and Eurasian kestrels to spatio-temporal variation of their

main prey. Ardea 97: 646-647.

Ahola, M., Laaksonen, T., Eeva, T., & Lehikoinen, E. 2009.
Great tits lay increasingly smaller clutches than selected for: a study of climate- and density-related changes in reproductive traits. Journal of Animal Ecology 78: 1298-1306.

Sirkiä, P.M., Laaksonen, T. 2009. Distinguishing between male and territory quality: females choose multiple traits in the pied flycatcher. Animal Behaviour 78: 1051-1060.

Lehtonen, P.K., Laaksonen, T., Artemyev, A.V., Belskii, E., Both, C., Bures, S., Bushuev, A.V., Krams, I., Moreno, J., Mägi, M., Nord, A., Potti, J., Ravussin, P.A., Sirkiä, P.M., Saetre, G.P. & Primmer, C.R. 2009, "Geographic patterns of genetic differentiation and plumage colour variation are different in the pied flycatcher (Ficedula hypoleuca)", Molecular Ecology 18: 4463-4476.

Lilley, T., Laaksonen, T., Huitu, O. & Helle, S. 2010. Maternal corticosterone but not testosterone level is associated with the ratio of second-to-fourth digit length (2D:4D) in field vole offspring (Microtus agrestis).
Physiology and Behaviour 99: 433-437.

Ruuskanen, S. & Laaksonen, T. 2010. Yolk androgens have sex-specific effects on behavior in the pied flycatcher Ficedula hypoleuca. Hormones and Behavior 57: 119-127.


Brommer, J., Alho, J.S., Biard, C., Chapman, J.R., Charmantier, A., Dreiss, A., Hartley, I.R., Hjernquist, M.B., Kempenaers, B., Komdeur, J., Laaksonen, T., Lehtonen, P.K., Lubjuhn, T., Patrick, S.C., Rosivall, B., van Oers, K., van der Velde, M., Tinbergen, J.M., Wilk, T., Winkel, W. 2010. Passerine extra-pair mating dynamics: A Bayesian modeling approach comparing four species. American Naturalist 176: 176-187.


Mäntylä, E., Sirkiä, P.M., Klemola, T., Laaksonen, T. 2010. Territory choice of pied flycatchers is not based on induced cues of herbivore-damaged trees. Open Ornithology 3: 105-111.


Sirkiä, P.M., Virolainen, M., Laaksonen, T. 2010. Melanin coloration has temperature-dependent effects on breeding performance that may maintain phenotypic variation in a passerine bird. Journal of Evolutionary Biology 23: 2385-2396.


Laaksonen, T., Adamczyck, F., Ahola, M.P., Möstl, E., Lessells, C.M. 2011. Yolk androgens and sexual conflict in the pied flycatcher. Behavioral Ecology and Sociobiology 65: 257-264.


Eeva, T., Ruuskanen, S., Salminen, J-P., Belskii, E., Järvinen, A., Kerimov, A., Korpimäki, E., Krams, I., Moreno, J., Morosinotto, C., Mänd, R., Orell, M., Qvarnström, A., Siitari, H., Slater, F.M., Tilgar, V., Visser, M.E., Winkel, W., Zang H., Laaksonen, T. 2011. Geographical trends in the yolk carotenoid composition of the pied flycatcher (Ficedula hypoleuca). Oecologia 165: 277-287.


Mäntylä, E., Klemola, T., Laaksonen, T. 2011. Birds help plants – a meta-analysis of trophic cascades caused by avian predators. Oecologia 165: 143-151.


Ruuskanen, S., Helle, S., Ahola, M., Adamczyck, F., Möstl, E., Laaksonen, T. 2011. Digit ratios have poor indicator value in a wild bird population. Behavioral Ecology and Sociobiology 65: 983-994.


Merilaita, S., Vallin, A., Kodandaramaiah, U., Dimitrova, M., Ruuskanen, S., Laaksonen, T. 2011. Number of eyespots and their intimidating effect on naïve predators in the peacock butterfly. Behavioral Ecology 22: 1326-1331.


Vasko, V., Laaksonen, T., Valkama, J., Korpimäki, E. 2011. Breeding dispersal of Eurasian kestrels under temporally fluctuating food abundance.  Journal of Avian Biology 42: 552-563.


Ruuskanen, S., Siitari, H., Eeva, T., Belskii, E., Järvinen, A., Kerimov, A., Krams, I., Moreno, J., Morosinotto, C., Mänd, R., Möstl, E., Orell, M., Qvarnström, A., Salminen, J-P., Slater, F.M., Tilgar, V.,  Visser, M.E., Winkel, W., Zang, H., Laaksonen, T. 2011. Geographical variation in egg mass and egg content in a passerine bird. PLoS ONE 6: e25360.


Ahola, M., Laaksonen, T., Eeva, T., Lehikoinen, E. 2012. Long-term changes and density-dependence in selection on timing of breeding and clutch size in the pied flycatcher. Oecologia 168: 703-710.


Santangeli, A., Hakkarainen, H., Laaksonen, T., Korpimäki, E. 2012. Home range size is determined by habitat composition but feeding rate by food availability in male Tengmalm’s owls. Animal Behaviour 83: 1115-1123.


Lehtonen, P. K., Laaksonen, T., Artemyev, A. V., Belskii, E., Berg, P. R., Both, C.,  Buggiotti, L., Bureš, S., Burgess, M. D., Bushuev, A.V., Krams, I., Moreno, J., Mägi, M., Nord, A., Potti, J., Ravussin, P-A., Sirkiä, P. M., Sætre, G-P., Winkel, W., Primmer, C. R. 2012. Candidate genes for colour and vision exhibit signals of selection across the pied flycatcher (Ficedula hypoleuca) breeding range. Heredity 108: 431-440.


Eeva, T., Andelmin, P., Hokkanen, T., Riivari, P., Ahola, M.P., Laaksonen, T., Lehikoinen, E. 2012. Breeding time trends of the crested tit (Lophophanes cristatus) in southern Finland - comparison of data sources. Journal of Ornithology 153(3): 653-661. DOI: 10.1007/s10336-011-0783-1


Burger, C., Belskii, E., Eeva, T., Laaksonen, T., Mägi, M., Mänd, R., Qvarnström, A., Slagsvold, T., Veen, T., Visser, M. 2012 Climate change, breeding date and nestling diet: how temperature differentially affects seasonal changes in pied flycatcher diet depending on habitat variation. Journal of Animal Ecology 81: 926-936. DOI: 10.1111/j.1365-2656.2012.01968.x 


Ruuskanen, S., Doligez, B., Pitala, N., Gustafsson. L., Laaksonen, T. 2012. Long-term fitness consequences of high yolk androgen levels: sons pay the costs. Functional Ecology 26: 884-894. DOI: 10.1111/j.1365-2435.2012.01994.x


Ruuskanen, S., Doligez, B., Gustafsson, L., Laaksonen, T. 2012. Long-term effects of yolk androgens on phenotype and parental feeding behaviour in a wild passerine. Behavioral Ecology and Sociobiology 66: 1201-1211. DOI: 10.1007/s00265-012-1373-9


Schuett, W., Laaksonen, J., Laaksonen, T. 2012. Prospecting at conspecific nests and exploration in a novel environment are associated with reproductive success in the jackdaw. Behavioral Ecology and Sociobiology 66: 1341-1350. DOI: 10.1007/s00265-012-1389-1


Santangeli, A., Lehtoranta, J., Laaksonen, T. 2012. Successful voluntary conservation of raptor nests under intensive forestry pressure in a boreal landscape. Animal Conservation 15: 571-578. DOI: 10.1111/j.1469-1795.2012.00551.x


Leskinen, P.K., Laaksonen, T., Ruuskanen, S., Primmer, C., Leder, E.H. 2012. The proteomics of feather development in pied flycatchers (Ficedula hypoleuca) with different plumage colouration. Molecular Ecology 21: 5762-5777. DOI: 10.1111/mec.12073


Helle, S., Laaksonen, T., Huitu, O. 2013. Sex-specific offspring growth according to maternal testosterone, corticosterone and glucose levels. Behavioral Ecology 24: 205-212. DOI:10.1093/beheco/ars155


Santangeli, A., Wistbacka, R., Hanski, I., Laaksonen, T. 2013. Ineffective enforced legislation for nature conservation: A case study with Siberian flying squirrel and forestry in a boreal landscape. Biological Conservation, 157: 237-244. DOI: 10.1016/j.biocon.2012.09.012.


Morales, J., Ruuskanen, S.,  Laaksonen, T., Mateo, R., Belskii, E., Eeva, T., Ivankina, E.V., Järvinen, A., Kerimov, A., Korpimäki, E., Krams, I., Mänd, R., Morosinotto, C., Orell, M., Qvarnström, A., Siitari, H., Slater, F.M., Tilgar, V., Visser, M.E., Winkel, W., Zang, H., Moreno, J. Variation in eggshell traits between geographically distant populations of pied flycatchers. Journal of Avian Biology 44:111-120.


Ruuskanen, S., Lehikoinen, E., Nikinmaa, M., Siitari, H., Waser, W., Laaksonen, T. Long-lasting effects of yolk androgens on phenotype in the pied flycatcher (Ficedula hypoleuca). Behavioral Ecology and Sociobiology, in press.


Santangeli, A., Högmander, J., Laaksonen, T. Returning white-tailed eagles breed as successfully in landscapes under intensive forestry regimes as in protected áreas. Animal Conservation, in press.


Sirkiä, P.M., Virolainen, M., Lehikoinen, E., Laaksonen, T. Fluctuating selection and immigration as determinants of the phenotypic composition of a population. Oecologia, in press.


Björklund, H., Valkama, J., Saurola, P., Laaksonen, T. Evaluation of artificial nests as a conservation tool for three forest-dwelling raptors. Animal Conservation, in press.


Ruuskanen, S., Laaksonen, T. Sex-specific effects of yolk androgens on begging behavior and digestion in pied flycatchers (Ficedula hypoleuca). Journal of Avian Biology, in press.


Thomson, R.L. Sirkiä, P.M., Villers, A., Laaksonen, T. Temporal peaks in social information: prospectors investigate conspecific nests after a simulated predator visit. Behavioral Ecology and Sociobiology, in press.