Peer-reviewed articles
2025
Kallio, P., Metsäranta, N. & Honkola, T. 2025. The language-contact landscape in the Baltics during the first millennium BC. Finnisch-Ugrische Forschungen. 70: 27–80. https://doi.org/10.33339/fuf.147292.
Metsäranta, N., Milanova, V. & Honkola, T. 2025. An Areal-Etymological Approach to Semantic Reconstruction of Proto-Uralic Kin Terms. Finnisch-Ugrische Forschungen. 70: 113–174. https://doi.org/10.33339/fuf.146092.
Nitch, A., Lummaa, V., Ketola, T., Honkola, T., Vesakoski, O. & Briga, M. 2025. The spatial distribution of pertussis, but not measles or smallpox, in pre-industrial Finland matches dialects. iScience. https://doi.org/10.1016/j.isci.2025.112530
Pesonen, P., Piha, M., Ahola, M., Holmqvist, E. & Honkola, T. 2025. Pottery, language and iron: reassessing Luukonsaari and Sirnihta Wares and their cultural context in Early Iron Age Finland. Documenta Praehistorica. 52: 2-27. https://doi.org/10.4312/dp.52.4
Santaharju, J., Syrjänen, K., Honkola, T., Seppä, P., Vesakoski, O. & Leino U. 2025. Data Release: Digitized Dialect Atlas of Finnish by Lauri Kettunen. Digital Humanities in the Nordic and Baltic Countries Publications. 7: 3. https://doi.org/10.5617/dhnbpub.12270.
2024
Milanova, V., Metsäranta, N. & Honkola, T. 2024. Kinship terminologies of the Circum-Baltic Area: Convergences and Structural Properties. Journal of Language Contact. 17: 315-358. https://doi.org/10.1163/19552629-bja10079
2023
Honkola, T., & Jordan, F. M. 2023. Kin Term Borrowings in the World’s Languages. Journal of Language Contact. 15: 562-626. https://doi.org/10.1163/19552629-15030004
Metsäranta, N., Milanova, V. & Honkola, T. 2023. Borrowability of kinship terms in Uralic languages. Finnisch-Ugrische Forschungen 68. https://doi.org/10.33339/fuf.120920
Moilanen, U., Salmela, E. & Honkola, T. 2023. Families on the move? The case of Proto-Finnic speakers. Moving northward: Professor Volker Heyd’s Festschrift as he turns 60. Lahelma, A., Lavento, M., Mannermaa, K., Ahola, M., Holmqvist, E. & Nordqvist, K. (eds.). [Helsinki]: p. 311-329, 19 p. (Monographs of the Archaeological Society of Finland; no. 11).
Passmore, S., Barth, W., Greenhill, S.J., Quinn, K., Sheard, C., Argyriou, P., Birchall, J., Bowern, C., Calladine, J., Deb, A., Diederen, A., Metsäranta, N.P., Araujo, L.H., Schembri, R., Hickey-Hall, J., Honkola, T., Mitchell, A., Poole, L., Rácz, P.M., Roberts, S.G., Ross, R.M., Thomas-Colquhoun, E., Evans, N., Jordan, F. M. 2023. Kinbank: A global database of kinship terminology. PLoS ONE 18(5): e0283218. https://doi.org/10.1371/journal.pone.0283218
2022
Norvik, M., Jing, Y., Dunn, M., Forkel, R., Honkola, T., Klumpp, G., Kowalik, R., Metslang, H., Pajusalu, K., Piha, M., Saar, E., Saarinen, S. & Vesakoski, O. 2022. Uralic typology in the light of a new comprehensive dataset. Journal of Uralic Linguistics. 1: 4-42. https://doi.org/10.1075/jul.00002.nor
Lynch, R., Loehr, J., Lummaa, V., Honkola, T., Pettay, J. & Vesakoski, O. 2022. Socio-cultural similarity with host population rather than ecological similarity predicts success and failure of human migrations. Proceedings of the Royal Society B. 289: 20212298. https://doi.org/10.1098/rspb.2021.2298
2021
Syrjänen, K.*, Maurits, L.*, Leino, U., Honkola, T., Rota, J. & Vesakoski, O. 2021. Crouching TIGER, hidden structure: Exploring the nature of linguistic data using TIGER values. Journal of Language Evolution. lzab004, https://doi.org/10.1093/jole/lzab004 *Shared first authorship.
Rantanen, T., Tolvanen, H., Honkola, T., & Vesakoski, O. 2021. A comprehensive spatial model for historical travel effort – a case study in Finland. Fennia – International Journal of Geography 199: 61–88. https://doi.org/10.11143/fennia.98357
Ketola, T., Briga, M., Honkola, T. & Lummaa, V. 2021. Town population size and structuring into villages and households drive infectious disease risks in pre-healthcare Finland. Proceedings of the Royal Society B. 288: 20210356. https://doi.org/10.1098/rspb.2021.0356
2019
Maurits, L., de Heer, M., Honkola, T., Dunn, M. & Vesakoski, O. 2019. Best practices in justifying calibrations for dating language families. Journal of Language Evolution. lzz009, https://doi.org/10.1093/jole/lzz009
Honkola, T., Syrjänen, K., Santaharju, J. & Pajusalu, K. 2019. Clustering lexical variation of Finnic languages based on Atlas Linguarum Fennicarum. Linguistica Uralica 55:3. https://dx.doi.org/10.3176/lu.2019.3.01
2018
Tambets, K., Yunusbayev, B., Hudjashov, G., Ilumäe, A-M., Rootsi, S., Honkola, T.,Vesakoski, O., Atkinson, Q. et al. 2018. Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations. Genome Biology. 19:139. https://doi.org/10.1186/s13059-018-1522-1.
Honkola, T., Ruokolainen, K., Syrjänen, K. J. J., Leino U, Tammi, I, Wahlberg, N. & Vesakoski, O. 2018. Evolution within a language: Environmental differences contribute to divergence of dialect groups. BMC Evolutionary Biology. 18:132. https://doi.org/10.1186/s12862-018-1238-6
2016
Syrjänen, K., Honkola, T., Lehtinen, J., Leino, A. & Vesakoski, O. 2016. Applying population genetic approaches within languages: Finnish dialects as linguistic populations. Language Dynamics and Change. 6: 235–283. DOI: 10.1163/22105832-00602002
2015
Ala-Honkola, O., Laine, L., Pekkala, N., Kotiaho, J. S., Honkola, T. & Puurtinen, M. 2015. Males Benefit from Mating with Outbred Females in Drosophila littoralis: Male Choice for Female Genetic Quality. Ethology 121: 577–585. DOI: 10.1111/eth.12369
2014
Lehtinen, J., Honkola, T., Korhonen, K., Syrjänen, K., Wahlberg, N. & Vesakoski, O. 2014. Behind family trees: Secondary connections in Uralic language networks. Language Dynamics and Change 4: 189-221.
2013
Honkola, T., Vesakoski, O., Korhonen, K., Lehtinen, J., Syrjänen, K. & Wahlberg, N. 2013. Cultural and climatic changes shape the evolutionary history of the Uralic languages. Journal of Evolutionary Biology 26: 1244–1253. DOI: 10.1111/jeb.12107.
Syrjänen, K., Honkola, T., Lehtinen, J., Korhonen, K., Vesakoski, O. & Wahlberg, N. 2013. Shedding more light on language classification using basic vocabularies and phylogenetic methods: A case study of Uralic. Diachronica 30: 323-352. DOI: 10.1075/dia.30.3.02syr.
Data releases
2024
Santaharju, J., Jordman, P., Honkola, T., Rantanen, T., Tammi, I., Koskela, S., & Vesakoski, O. 2024: A Digitized Historical Database of Cultural, Economic, Demographic, and Ecological Data of Finland (CEDEDA) (0.1) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.13975170
Santaharju, J., Syrjänen, K., Honkola, T., Seppä, P., Vesakoski, O. & Leino, U. 2024. New version of the digitized Dialect Atlas of Finnish by Lauri Kettunen [Data set]. Zenodo. https://doi.org/10.5281/zenodo.10078078
2022
Passmore, S., Sheard, C., Argyriou, P., Bowern, C., Calladine, J., Deb, A., Diederen, A., Hickey-Hall, J., Honkola, T., Mitchell, A., Poole, L., Rácz, P., Roberts, S., Ross, R., Thomas-Colquhoun, E. & Jordan, F. M. 2022. Varikin: A global collection of kinship terminology [Data set]. Zenodo. https://doi.org/10.5281/zenodo.6562609 https://github.com/kinbank/varikin
Honkola, T., Metsäranta, N., Milanova, V., Passmore, S., & Jordan, F. M. 2022. Kinura: A database of kinship terminology from the Uralic Language family [Data set]. Zenodo. https://doi.org/10.5281/zenodo.6673055 https://github.com/kinbank/kinura
Norvik, M., Jing, Y., Dunn, M., Forkel, R., Honkola, T., Klumpp, G., Kowalik, R., Metslang, H., Pajusalu, K., Piha, M., Saar, E., Saarinen, S. & Vesakoski, O. 2022. Uralic typology in the light of a new comprehensive dataset [Data set]. Zenodo. DOI:10.5281/zenodo.5236365.
Non-peer reviewed articles
Honkola, T. 2016. Kielten makro- ja mikroevoluutio: Kielten erkaantumisen tutkiminen evoluutiobiologisista lähtökohdista. [Macro- and microevolution of languages: Exploring linguistic divergence with approaches from evolutionary biology]. Lectio praecursoria. Online version of the journal Virittäjä 1.
Publications intended for general public
Tambets, K., Honkola, T. & Metspalu, M. Whole genome-wide study reveals most Uralic-speakers have traces of recent common demographic history in their genes. On Biology Blog 24.10.2018
Honkola, T. Using a biological framework to resolve the early stages of linguistic divergence BMC Series Blog 6.9.2018.
Theses
Honkola, T. 2016. Macro- and microevolution of languages: Exploring linguistic divergence with approaches from evolutionary biology. PhD thesis. University of Turku, Turku.
Honkola, T. 2009. The effects of inbreeding, crossbreeding and stress on metabolic rate in Drosophila littoralis. MSc thesis. University of Jyväskylä.
Honkola, T. 2007. Takasiiven värityksen ja melanismikuvion periytyminen täpläsiilikäskoiraalla (Parasemia plantaginis). [The inheritance of hindwing color and melanin pattern in wood tiger moth males (Parasemia plantaginis)] BSc thesis. University of Jyväskylä.