Morphology and systematics of moths and butterflies

Past research

During the 1980s and 1990s, I intensively studied the thoracal morphology of moths and butterflies in connection with insect flight. I described over 100 taxa of fossil and recent insects (primarily moths), ranging from sub-species to sub-orders. My taxonomic research focussed on archaic moths of the families Micropterigidae, Eriocraniidae, Opostegidae, Tisheriidae, Prodoxidae, Incurvariidae and Adelidae.

Ongoing research

I am preparing for publication descriptions of over 100 new species of the genus Nemophora (Lepidoptera: Adelidae) and am analysing the phylogeny of the family Adelidae based on morphological and DNA data.

Featured publications

Kozlov, M. V. (2016) Taxonomic revision of Australian long-horn moths of the genus Nemophora (Lepidoptera: Adelidae). Zootaxa4097, 84–100 (doi: 10.11646/zootaxa.4097.1.4)

The fauna of Australia currently includes seven valid species of Nemophora, which form a monophyletic laurella species-group. Keys to species based on external characters and on male genitalia are provided; adults and male genitalia are described and illustrated.

Kozlov, M. V. (2013) A new species of the genus Adela (Lepidoptera: Adelidae) from South America. Neotropical Entomology42, 505-507 (doi: 10.1007/s13744-013-0150-4).

Adela boliviella n. sp., described from central Bolivia, is the first representative of the subfamily Adelinae found in South America.

Kozlov M. V. (2011) Ground plan and evolution of pterothoracic musculature of moths and butterflies (Lepidoptera). Entomological Review90, 833-853 (doi: 10.1134/S0013873812020054)

Investigation of the meso- and metathoracic musculature in 62 species of Lepidoptera revealed 40 topographically different muscles, 37 of which presumably belong to the ground plan. The thoracic anatomy of Lepidoptera is strongly linked with biomechanics of flight and therefore has a relatively low value for phylogenetic analysis.

Kozlov, M. V. (2004) Annotated checklist of the European species of Nemophora (Adelidae). Nota Lepidopterologica26, 115-126. (PDF)

The checklist includes 20 species so far reported from Europe, along with most important synonymy and comments about the characters useful for identification of the problematic species.


D. Agassiz
G. Esartia
D. Grodnitsky
T. Hirovatari
P. Huemer
V. Ivanov
L. Kaila
O. Karsholt
†N. P. Kristensen
†V. I. Kuznetzov
M. Motorkin
M. Mutanen
M. Nesina
E. J. van Nieukerken
R. Pupljasis
A. P. Rasnitsyn
†G. S. Robinson
S. Seksyaeva
S. Sinev
A. A. Stekolnikov
N. Wahlberg

Co-operating scientists

D. Davis
W. Mey
L. Przybylowicz
K. Tuck
S.-H. Yen
and over 50 other taxonomists from around the world


EC through SYS Resource and SYNTHESYS programmes
Academy of Finland
Finnish Cultural Foudation
Oskar Öflunds Stiftelse